unaware of any study that has examined the in?uence of thin layers on the behavior of individual gelatinous Spatial and temporal heterogeneity affect movement zooplankton. and resource use by animals (Levin et al. , 2000). In Gelatinous zooplankton, including ctenophores, particular, behaviors that allow organisms to locate medusae, and siphonophores, tend to aggregate at and exploit patches of food represent key in?uences density, salinity, and temperature discontinuities that on trophodynamics. For plankton, external forcing develop in coastal transition zones, estuaries, and (e. g. , currents, shear, and turbulence) combines with fjords (Owen, 1989; Graham et al. , 2001). Aggre- behavioral responses to determine the structure and