This work provides in-depth analysis of the origins of landscape ecology and its close alignment with the understanding of scale, the causes of landscape pattern, and the interactions of spatial pattern with a variety of ecological processes. The text covers the quantitative approaches that are applied widely in landscape studies, with emphasis on their appropriate use and interpretation. The field of landscape ecology has grown rapidly during this period, its concepts and methods have matured, and the published literature has increased exponentially. Landscape research has enhanced understanding of the causes and consequences of spatial heterogeneity and how these vary with scale, and they have influenced the